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Featured California Wildflowers

Sierra Nevada Wildflowers, Saddlebag Lake, California.

*STEER'S HEAD AND ITS COUSIN

One look at a Steer's Head flower and you will never forget the plant's name -- the most appropriately named native plant in the Sierra Nevada Mountains.


*ELEPHANT'S HEAD AND LITTLE ELEPHANT'S HEAD

Other appropriately named Sierra Nevada wildflowers.

*HYBRID COLUMBINES

A beautiful hybrid found only in the high altitudes of the Sierra Nevada.

*SNOW PLANT

The most popular flowering plant in the Sierra Nevada, according to John Muir.

*OTHER NON-PHOTOSYNTHETIC PLANTS

An introduction to parasitic plants found in the Sierra Nevada, including several examples.

*Plant Parasite Conundrum

Three species of wintergreen (genus Pyrola) exhibit varying degrees of parasitism.

*SIERRA NEVADA ORCHIDS

A presentation of several Sierra Nevada orchid species.

*Gentians

Images and information of gentian species found in the Sierra Nevada.

*America's Peonies

The only two native peonies of the United States both grow in California.

*SIERRA'S SHOOTING STARS

It's easy to recognize a Shooting Star; how do you distinguish among the species?

steer's head and its cousin

The most appropriately named native plants.

California Wildflower: Steer's head, Dicentra uniflora. Sonora Pass California.

steer's head Flower

Some plants are unforgettable because they are appropriately named. Steer's Head (Dicentra uniflora) is one such plant. The plant's pinkish-cream-colored flower resembles the skull of a Texas longhorn.

Steer's head's, Dicentra uniflora, size in relation to US quarter. Yosemite NP, California.

Steer's head size in relation to U.S. quarter

Steer's Head is one of the earliest bloomers.  It is often found just a few feet from the edge of a melting snowbank. Steer's Head may be overlooked—the color of its tiny flower blends in with rocky soils covered by pine needles. The plant is most easily spotted by searching for its grayish-blue-green divided leaf.

Steer's head, Dicentra uniflora, flower and leaf. Near Sonora Pass, California.

Steer's Head flower and leaf stems attach underground

The flower and leaf stems of the Steer's Head attach below ground and sometimes do not look like they belong to the same plant. Look around once you find a plant. There will likely be more.

Steer's Head, Dicentra uniflora, California wildflower playing card.

Steer's head card, deck a

Pacific Bleeding Heart, Dicentra formosa, California wildflower playing card.

Pacific Bleeding heart flower

Steer's Head's cousin,  Pacific Bleeding Heart (Dicentra formosa), also has a fitting name.  The heart-shaped flowers of this plant may not immediately resemble the Steer's Head flower. Imagine pulling back the spreading tips of the Bleeding Heart's petals. They will resemble the "horns" of the Steer Head's flower. 

Pacific bleeding heart flower and leaves

Pacific Bleeding Heart and Steer's Head plants are part of the poppy family, Papaveraceae.  Characteristic of the poppy family, both plants have highly divided, fern-like, blue-green leaves. 


Pacific Bleeding Heart can be found in moist, shaded locations.

pacific bleeding heart carD, deck b


Elephant's head and Little Elephant's head

Runners-up for the most appropriately named plants.

Sierra Nevada wildflower identification photograph: Elephant's Head, Pedicularis groenlandica.

Elephant's head

The Elephant's Head (Pedicularis groenlandica) of the paintbrush family, Orobanchaceae, is another aptly named and unforgettable plant. It grows from one to three feet tall and has a stem containing numerous small flowers (an inflorescence). Each flower resembles an elephant's head with floppy ears and a trunk. Elephant's Head grows in wet meadows and along streams.

Elephant's Head, Pedicularis groenlandica, California wildflower playing card.

Elephant's Head card, Deck A

California Wildflower: Little elephant's head, Pedicularis attollens. Dana Pass trail, California.

Little Elephant's head

A related plant of Elephant's Head, Little Elephant's Head (Pedicularis attollens), has smaller flowers and a more cartoonish elephant face with distinct eye spots. It inhabits similar environments as the larger species but has a later bloom season.

Little Elephant's Head, Pedicularis attollens, California wildflower playing card.

Little Elephant's head card, deck b


Hybrid columbines

California Wildflower: Crimson columbine, Aquilegia formosa, flower. Tuolumne County, California.

crimson columbine

Crimson Columbine (Aquilegia formosa) is common in the Sierra Mountains and California's coastal regions.  It grows from one to three feet in partial shade of moist forested areas, meadows, and along stream banks. The leafy portion of the plant forms a mound.  Long branched stems rise above the mound and produce several beautiful nodding flowers with red to red-orange and yellow pigmentation.  Native Americans used this plant for several medicinal purposes.  Caution: columbine plants are toxic.


Crimson Columbine, Aquilegia formosa, California wildflower playing card.

crimson columbine card, deck a

California Wildflower: Sierra columbine, Aquilegia pubescens, flower.  Mount Dana Pass trail, California.

Sierra Columbine

Sierra Columbine (Aquilegia pubescens) is a California endemic species limited to subalpine forests and alpine fell fields of the Sierra Nevada Mountains.  Sierra Columbine's flowers are larger than the flowers of Crimson Columbine and are more erect  (less nodding). Flowers are white, cream, or yellow with tinges of pink.  

Sierra Columbine, Aquilegia pubescens, California wildflower playing card.

Sierra Columbine card, deck b

Moth pollinating columbine flower.  Saddlebag Lake, California.

hybrid columbine & moth pollinator

Hummingbirds pollinate the lower elevation Crimson Columbine, while moths pollinate the higher elevation Sierra Columbine. In areas where the ranges of the two species overlap, the same animal may pollinate both species. This sometimes results in stunningly beautiful hybrid plants with larger Sierra Columbine flowers and the expression of color genes from Crimson Columbine.

California wildflower landscape photograph. Aquilegia formosa X Aquilegia pubescens hybrid.

Aquilegia formosa X Aquilegia pubescens HYBRID


Snow Plant

Arguably the most popular flowering plant in the Sierras.

Snow Plant in forest. Lily Lake, Tuolumne County, California.

Snow plant

In his book The Yosemite (1912), John Muir wrote, “The Snow Plant  (Sarcodes sanguinea) is more admired by tourists than any other in California.”  Visitors have continued to be intrigued by this plant for the 110+ years since Muir's statement. Snow Plants are among the most recognizable plants in California. They are easy to locate and identify as their bright red bodies contrast strikingly against the brown hues of the forest floor.  

Snow Plant, Sarcodes sanguinea, in snow.  Tuolumne County, California.

Young snow plant after spring storm

The Snow Plant obtained its name from the perception that it grows through melting snow. This rarely happens. It is more likely for an already emerging plant to be covered with the snow from a late spring storm. 

Sierra Nevada wildflower identification photograph. Snow Plant, Sarcodes sanguinea.

Stalk, Bracts, and Flowers of snow plant

Snow Plants generally grow to about a foot in height. The above-ground portion consists of a thick stalk supporting multiple flowers and bracts: the large size and bright color aid pollinators in locating snow plants. 

Young snow plant, Sarcodes sanguinea, stalks starting to grow. Tuolumne County, California.

young flower stalks as seen from above

Snow Plants lack the green pigment chlorophyll and do not perform photosynthesis. Instead, they obtain carbon compounds from soil fungi (mycorrhizae). The soil fungi have a mutually beneficial relationship with conifers. The fungi enhance the tree's uptake of water and nutrients. In return, the trees provide the fungi with carbon compounds. The carbon is then shuttled to the Snow Plants, but Snow Plants provide the fungi nothing in return. Thus, the Snow Plant is a parasite of the fungus.

Snow Plant, Sarcodes sanguinea, California wildflower playing card.

Snow Plant Card. DECK A


Other non-photosynthetic plants

Lesser known parasitic plants.

California wildflower identification photograph. Pine Drops, Pterospora andromedea.

recognizing parasitic plants

You may be surprised to learn several entirely parasitic plants exist in the Sierra Nevada Mountains. They are less well-known than Snow Plants because they are smaller, not as brightly pigmented, or less common. They all share a lack of chlorophyll, the pigment that makes plants green.  Parasitic plants are reddish, yellow, purple, or white. They lack or have reduced leaves, consisting mainly of roots and stems supporting flowers.  A few examples of parasitic plants follow.

Pine drops,  Pterospora andromedea. Near Eagle Meadow, Tuolumne County, California.

pine drops

Pine Drops (Pterospora andromedea) are common but often overlooked parasitic plants in the understory of California's yellow pine and red fir forests.  From a distance, plants may look like two to three-foot knobby sticks poking out of the soil. These are the plants' stems and flowers. Like Snow Plants, Pine Drops are non-photosynthetic plants and rely on soil fungi for survival.   

Pine Drops, Pterospora andromedea, California wildflower playing card.

pine drops card, deck b

California wildflower identification photograph. Striped Coralroot, Corallorhiza striata.

Young striped coralroot plant

Striped Coralroot (Corallorhiza striata) and the two following plants (Spotted Coralroot and Phantom Orchid) are parasitic orchids, part of the forest floor vegetation, in California's Sierra and coastal mountains. All three plants grow in decomposing leaf litter where the fungi they parasitize are abundant. Like Snow Plants and Pine Drops, fungi function as intermediaries in carbon transfer from nearby trees to these parasitic orchids.

Sierra Nevada wildflower identification photograph. Striped Coralroot, Corallorhiza striata.

Striped Coralroot Orchid

The above-ground portions of the Striped Coralroot Orchid consist of one or several flower stems containing a few to several flowers and sheath-like leaves wrapped around the base of each stem. The plant may be yellowish, salmon, or pinkish-purple. Striped Coralroot has distinct stripes on the sepals and petals. The lower lip petal generally has wider stripes.

Striped Coralroot, Corallorhiza striata, California wildflower playing card.

striped coralroot card, deck b

Spotted Coralroot Orchid

Spotted Coralroot (Corallorhiza maculata) may appear as a single stem or multiple stems arising from a rhizome. Like striped coralroot, the leaves are reduced to sheaths surrounding the stem. All parts of Spotted Coral root are reddish/brownish or yellowish in coloration, except for the lip petal, which is white with various degrees of spotting. 

Sierra Nevada wildflower identification photograph. Spotted Coralroot, Corallorhiza maculata,

Spotted Coralroot Flowers

Spotted Coralroot, Corallorhiza maculata, California wildflower playing card.

Spotted coralroot card, deck a

Numerous Phantom orchids growing in a forest

Spotting an elusive Phantom Orchid (Cephalanthera austiniae) is always a treat. It is found in the shade of an open forest, often on slopes and near bodies of water. The plant is white except for yellow on the flower's lower lip.  

Sierra Nevada wildflower identification photograph. Phantom Orchid, Cephalanthera austiniae.

Phantom orchid

Phantom Orchid, Cephalanthera austiniae, California wildflower playing card.

phantom orchid cards, deck b

Sierra Nevada wildflower identification photograph. Flat-topped broomrape, Aphyllon corymbosum

Flat-topped bromrape

Unlike the other parasitic plants described here, broomrape plants parasitize plant roots directly. Several broomrape species reside in California. Flat-topped Broomrape (Aphyllon corymbosum) occurs primarily in the Eastern Sierra, where it is a parasite of Sagebrush (Artemisia tridentata) roots. Flowers are a pale grayish or yellowish pink outside, making them difficult to spot against rocky substrate. They are deeper pink inside with dark veins. 

Flat-topped Broomrape, Aphyllon corymbosum, California wildflower playing card.

flat-topped broomrape cards, deck b

Clustered bromrape

Clustered Broomrape has been classified into two separate species: Aphyllon fasciculatum and Aphyllon franciscanum. Both species are found in the Sierra Nevada and other western and central United States regions. They share the characteristic of numerous flower stalks of pale yellow to reddish-pink. The mature flowers of A. fasciculatum are about the same height, and the petal lobe tips are rounded.  A. franciscanum has mature flowers of various heights with pointed petal lobes. 

Habitat of Purple Broomrape

A much smaller Purple or Naked Broomrape (Aphyllon purpureum) is also found in the Sierra Nevada Mountains. Purple Broomrape is a two-inch, single-stalked plant that parasitizes plants of the stonecrop, sunflower, or saxifrage families. The photo above shows Broadleaf Stonecrop (Sedum spathulifolium) growing on a moist, shaded rock ledge. The small Purple Broomrape (insert and arrows) grows almost unnoticed, among sedum leaves, as it steals nutrients from the host plant’s roots.

Purple Broomrape Flower

The purple or white petals of Purple Broomrape are fused to form a long corolla tube. The petals are darkly veined. The lower petal folds may be yellow to yellow-orange. Petals, sepals, and the stalk are covered with minute glandular hairs. 

California wildflower identification Photograph. Canyon Dodder, Cuscuta subinclusa.

Canyon dodder on host plant

Dodder plants employ an entirely different parasitic strategy. The Canyon Dodder (Cuscuta subinclusa) has golden stems that look like string. These stems blanket the host plant and twine around the host plant’s stems. At points where the stems of the parasite and host are in contact, specialized roots known as haustoria grow from the parasite’s stem to penetrate the host plant. Dodder obtains water and nutrients from the host’s vascular tissue via the haustoria. 

Sierra Nevada wildflower identification photograph. Canyon Dodder, Cuscuta subinclusa.

canyon dodder twining around host stem; dodder flowers

Canyon Dodder produces clusters of tiny, white, bell-shaped flowers. The seeds germinate in the soil and have only days to locate, grow towards, and attach to an appropriate host, or they will die. The seedlings identify host plants by detecting chemicals the host plant releases into the air.  


Plant Parasite Conundrum

The Pyrola Pica Species Complex

Typical habitat for the Pyrola Pica Species Complex

All non-photosynthetic, parasitic plants descended from photosynthetic ancestors. Initially relying on fungi or other plants for enhanced water and nutrient absorption, they gradually became dependent on their hosts for carbon compounds. Here, three closely related species of wintergreen (family Ericaceae), including both fully parasitic and partially parasitic members, are discussed to gain insight into the origins of plant parasitism.

California wildflower. White-veined Wintergreen, Pyrola pica, viewed from above.

Distinctive leaf markings of White-veined Wintergreen

White-veined Wintergreen (Pyrola pica), Toothed Wintergreen (P. dentata), and Leafless Wintergreen (P. aphylla) sometimes grow together in the thick litter layer of Sierra Nevada forests, where they occasionally hybridize. They share many morphological features and are all associated with soil fungi. Botanists have debated whether these plants comprise a single species or are three separate species. Today, genetic evidence supports the view that they are three distinct, closely related species. 

Sierra Nevada photography. White-veined Wintergreen, Pyrola pica.

White-veined Wintergreen’s rosette of leaves

White-veined Wintergreen is easily identifiable by its dark green, white-veined rosette of spreading leaves. White-veined Wintergreen performs photosynthesis, yet receives some of its carbohydrates from soil fungi. Occasionally, a normally leaved plant and a stalk with significantly reduced leaves (more parasitic) will originate from the same rhizome, revealing the nutritional benefits derived from White-veined Wintergreen’s interaction with soil fungi and the plant’s semi-parasitic lifestyle.

Flower structure of White-veined Wintergreen

White-veined Wintergreen flowers are representative of the flower structure of all Pyrola species. Few to many nodding flowers are arranged on an unbranched stem. A small bract is located at the base of each flower stalk. Small sepals surround fleshy, white, pink, or green, roundish petals. Ten stamens with two-chambered anthers have pore openings at the top, allowing pollen to be released. The single pistil has a long, often curved style that extends past the petals.

Toothed Wintergreen

Toothed Wintergreen resembles White-veined Wintergreen. The most apparent distinction is that Toothed Wintergreen has gray-green or blue-green erect leaves with less distinct vein markings and a powdery surface appearance, much like the powdery appearance of grape skin. Contrary to the plant’s name, its leaf margin may or may not be toothed. Like White-veined Wintergreen, Toothed Wintergreen may produce stalks with reduced leaves and relies on soil fungi for part of its nutritional needs.

California wildflower. Toothed Wintergreen, Pyrola dentata, viewed from above.

Toothed Wintergreen as seen from above

California wildflower identification photograph. Leafless Wintergreen, Pyrola aphylla.

Leafless Wintergreen Plant

Unlike the previous two wintergreen species, Leafless Wintergreen is entirely parasitic, having no chlorophyll and characterized by white or red coloration. Leafless Wintergreen appears leafless, but it produces small leaves near the base of the stem (lower photo insert), remnants of its photosynthetic ancestors' leaves. The litter layer often hides these vestigial leaves.

Sierra Nevada wildflower. Leafless Wintergreen, P.yrola aphylla, flowers.

Flowers of Leafless Wintergreen

The flower stems of Leafless Wintergreen can reach up to two feet in height and may occur singly or in groups. Stalks growing individually may be difficult to spot until a break in the forest's canopy spotlights the plant. From a distance, they may look like nothing more than upright twigs. Close up, each flower spike supports the same captivating flowers as the other wintergreen plants.


Sierra nevada orchids

Ecological associations hidden in plain sight.

Small flowers of a California orchid.

the small flowers of many orchids are easily overlooked

When most of us think of orchids, we envision the showy tropical epiphytic (growing on other plants) orchids sold in nurseries or used for corsages. California’s orchids, in contrast, are terrestrial, primarily small, not particularly colorful, and often overlooked. Although inconspicuous, California orchids abound with interspecies interactions hidden in plain sight.

Spotted Coralroot and Phantom Orchid growing together.

Phantom and Spotted coralroot orchids

Orchids have tiny seeds containing little nutrition. Consequently, orchid seeds require symbiotic relationships with soil fungi to obtain the nutrition needed for seed germination. As with the non-photosynthetic orchids addressed in a previous section, some photosynthesizing orchids maintain these fungal associations throughout their lives. Orchids also have complex, highly specific, and sometimes deceptive relationships with pollinators. 

California wildflower identification photograph. White Bog Orchid, Platanthera dilatata.

white bog orchid - Plathanthera species

Plathanthera is the largest orchid genus in North America. Plathanthera species have lip petals that curl in front and extend back to form a sac-like spur. The length of each species’ spur corresponds to the length of its pollinator’s (moth or butterfly) tongue. 

White Flowered Bog-Orchid, Platanthera dilatata, flower. Tuolumne County, California.

white bog orchid flower

Because of its height, numerous flowers, long flowering period, and wide distribution, the White Bog Orchid (Platanthera dilatata) is the orchid you will most likely encounter and recognize in the Sierras. White Bog Orchids always have white flowers and spurs longer than the lips. They grow in seeps, meadows, and along streams in full sun or partial shade.

White Bog Orchid, Platanthera dilatata, California wildflower playing card.

white bog orchid card, deck b

California Wildflower: Sparse flowered bog orchid, Platanthera sparsiflora. Herring Creek, Tuolumne County, California.

sparsely-flowered bog orchid

The Sparsely-flowered Bog Orchid (Platanthera sparsiflora) looks much like the White Bog Orchid and grows in similar habitats. Green flowers of Sparsely-flowered Bog Orchids distinguish them from similar plants. The flower spurs are about the same length as the lips.

California wildflower: Dense-flowered Rein Orchid, Piperia elongata

dense-flowered rein  orchid

The Dense-flowered Rein Orchid (Piperia elongata) is another green orchid that may look like the Sparse-flowered Bog Orchid to the casual observer. The Dense-flowered Rein Orchid has shorter sepals and petals and a longer spur. The plant has only basal leaves, lacking leaves on the stem. This rein orchid grows in dry forests, whereas the Sparse-flowered Bog Orchid is found in wet environments.

California wildflower identification photograph. Royal Rein Orchid, Piperia transversa.

Royal rein orchid

Royal Rein Orchids (Piperia transversa) are also found in dry woodlands. The plants have two leaves at their base, which wither before flowering to give the appearance of leafless flower stems. Royal Rein Orchids have white flowers with almost horizontal spurs that are much longer than the lips.

Royal Rein Orchid, Piperia transversa, California wildflower playing card.

ROYAL REIN ORCHID card, deck A

California wildflower identification photograph. Coleman’s Rein Orchid, Piperia colemanii.

Coleman’s Rein Orchid

Coleman’s Rein Orchid (Piperia colemanii) is a California endemic species with populations occurring within an elevation band of 4,000-6,500 feet from Siskiyou County to Fresno County. The plant produces a stem 6-21 inches tall with narrow (<0.5 inch width), grass-like leaves at the base. Numerous tiny green flowers require a magnifying glass to appreciate. The Flower’s lip-petal is longer than its spur.




California Wildflower: Hooded ladies tresses, Spiranthes romanzoffiana.  Mono County, California.

Hooded ladies' tresses

Charles Darwin first studied and described the pollination mechanism of Spiranthes species, such as Hooded Ladies' Tresses (Spiranthes romanzoffiana). Young flowers only partly open, preventing bee pollinators from accessing the nectary. As bees probe for nectar, they pick up pollen. Older flowers open wider, allowing the pollinator to reach the nectary. Pollen from the previous flower is transferred to the sticky stigma, promoting cross-pollination.

California wildflower identification photograph. Rattlesnake Plantain, Goodyera oblongifolia.

Rattlesnake Plantain

Rattlesnake Plantain (Goodyera oblongifolia) is found in full to partial shade in thick humus of coniferous forests at low and mid-elevations. Even without flowers, Rattlesnake Plantain is easy to recognize from its rosette of evergreen white-veined leaves. The leaf markings are highly variable. Generally, the midrib is white, but markings may be more extensive and quite beautiful.

Sierra Nevada photograph. Rattlesnake Plantain, Goodyera oblongifolia, flowers.

Rattlesnake Plantain Flowers

A young Rattlesnake Plantain plant may take several years to bloom. When it does, the buds mature slowly and will not open until late summer. The flower petals are white and slightly translucent. The sepals are white with tinges of green. The Rattlesnake Plantain is pollinated by bumblebees, utilizing the same pollination mechanism as Hooded Ladies' Tresses.

California wildflower identification photograph. Mountain Lady’s Slipper, Cypripedium montanum.

Mountain lady’s slipper plants

The distribution of Mountain Lady’s Slipper (Cypripedium montanum)  extends from Southern Alaska to the Central Sierra Nevada Mountains. Mountain Lady’s slipper may grow to over two feet in height under varied light and moisture conditions.

Sierra Nevada photograph. Mountain Lady’s Slipper, Cypripedium montanum.

Mountain lady’s slipper

The flower of the Mountain lady’s Slipper consists of three long, narrow, brownish-red sepals and two similar appearing twisted petals. A third petal forms a white pouch. Inside the white petal pouch are reddish nectar guides and scented hairs. 

 Mountain Lady’s Slipper, Cypripedium montanum. flower close up.

Mountain lady’s slipper flower

The nectar guides and hairs attract and direct insect pollinators, flies, or solitary bees into the pouch. Pollinators are led through a one-way passage where sticky pollen sacs attach to them. Ideally, pollen is transferred to the next flower visited by the insects. Pollinators are deceived and lured into the flower but receive no nectar reward.

 Mountain Lady’s Slipper, Cypripedium montanum,  wildflower playing card.

MOUNTAIN LADY’S SLIPPER cards, deck b

California wildflower identification photograph. Stream Orchid, Epipactis gigantea.

Stream Orchid plant

The Stream Orchid or Chatterbox (Epipactis gigantea) is another large orchid in the Sierra Nevada Mountains. It may grow over three feet tall and contain numerous flowers per stem. Stream Orchids can reproduce via rhizomes, sometimes resulting in large cloned populations.  

Sierra Nevada wildflower. Stream Orchid, Epipactis gigantea.

Stream Orchid

Stream Orchids grow only where their roots receive a continuous water supply. Although the Stream Orchid plant is large, the flowers are often less than two inches wide and may be difficult to spot among the dense foliage of its wet environment. However, the plant’s flowers are impressive, with green, pink, and yellow at close inspection. 

Sierra Nevada photography. Stream Orchid, Epipactis gigantea.

Stream Orchid flowers

Like the mountain lady’s slipper, the stream orchid deceives its pollinator. In this case, the flies pollinating stream orchids typically lay eggs in aphid nests. The aphids become food for the fly’s growing larvae. The scent of the stream orchid flower mimics that of aphids, prompting flies to lay eggs in stream orchid flowers. No aphids are present on the flower, so the hatched fly larvae die. The orchid benefits as adult flies transfer pollen sacs from one flower to another.

Stream Orchid, Epipactis gigantea, California wildflower playing card.

STREAM ORCHID cards, deck A


Gentians

From giants to dwarfs.

Centaury Plant belongs to the gentian family

The Flower Symmetry section of the Flower Morphology page introduced the Centaury Plant of the gentian family Gentianaceae. Other plants from this family may surprise you as they appear much different than the Centaury Plant and each other. With exceptions, gentian family plants have opposite leaves with smooth edges that lack leaf stalks. Petals overlap and twist when in the bud. Sepals, petals, and stamens are four or five in number. Sepals and petals are fused.  

 Sierra Nevada photography. Monument Plant, Frasera speciosa.

Several blooming Monument Plants

Due to its large size, the Monument Plant (Frasera speciosa) of the gentian family is one of the most recognizable plants in the Sierra. It is a perennial that can live for decades before blooming. For most of its life, the above-ground plant consists of a large rosette of leaves that die back each winter. As the plant prepares to produce flowers, it grows a stalk from two to over seven feet tall that towers over other plants in its meadow and open woodland habitats.  

 California wildflower identification photograph. Monument Plant, Frasera speciosa.

Monument Plant flowers and buds

The Monument Plant flowers only once before dying, but it does so with splendor, producing hundreds of flowers. Blooming is sporadic yet synchronized with other nearby but spatially separated plants of the same species. This strategy may deter herbivores who cannot rely on predictable flowering. The Monument Plant utilizes various pollinators, assuring some are available and pollination rates are high in the erratically timed flowering years.

Monument Plant, Frasera speciosa flower detail.

Monument Plant flower

Typically, each Monument Plant flower has long pointed sepals, pale cream-yellow-green petals having purple spotting, and two fringed nectaries. The anthers are large, and the central pistil is prominent. Collectively, these features create an unusual and impressive flower.

 Monument Plant, Frasera speciosa, California wildflowers playing cards.

Monument Plant playing card, deck A

California wildflower identification photograph. Kern Frasera, Frasera tubulosa.

Kern Frasera plant

California is home to eight Frasera species and several varieties. Kern Frasera (Frasera tubulosa) is an endemic southern Sierra Nevada species with limited distribution. The plant can grow to over three feet or may be much smaller, a dwarf compared to its giant cousin the Monument Plant. Leaves are linear and whorl around the base of the stem. Flowers are white to pale blue with darker blue veins. The plant lives in open conifer forests between 6,000 and 9,000 feet.

Kern Frasera, Frasera tubulosa, California wildflowers playing card.

Kern Frasera playing card, Deck B

Population of Sierra Alpine Gentian, Gentiana newberry, with flower close up.

Population of Sierra Alpine Gentian

The next four gentians are all late summer bloomers that grow in wet meadow environments. Pictured here is a population of Sierra Alpine Gentian (Gentiana newberry). Each plant has a rosette of leaves just above ground level. The stem attaches below these leaves. It grows horizontally at first and then turns upward, having additional stem leaves and terminating with a disproportionately large flower to a height of about four inches. Each plant usually has one, but up to five flowers. 

Sierra Alpine Gentian plant

Sierra Alpine Gentian plant (insert: pale blue flower)

The flowers of Sierra Alpine Gentian have white or blue fused petals. Jagged, multiple-tipped accessory structures between each petal lobe are pigmented to match the inside petal. Petal lobes have green spots inside. Smaller green spots form the appearance of green lines further down the corolla tube. The exterior of the petals is marked with brown-maroon bands, creating a candy-stripe appearance.  

Sierra Nevada Photography. Sierra Alpine Gentian, Gentiana newberry, flower detail.

Sierra Alpine Gentian Flower

Two varieties of Sierra Alpine Gentian are found in the Sierra. Gentiana newberry tiogana is the smaller variety with white or pale blue flowers. A less common Gentiana newberry newberry, is the larger variety, with darker blue flowers.

Sierra Alpine Gentian, Gentiana newberry, California wildflowers playing cards.

Sierra Alpine Gentian card, Deck A

California wildflower identification photograph. Hiker's Fringed Gentian, Gentianopsis simplex.

Hiker's Fringed Gentian

The gentian family is known for including species with the most genuine blue colors. Hiker's Fringed Gentian (Gentianopsis simplex) has pale or intense, true-blue or purple flowers. The blue fringed petals contrast with the white throat of the corolla. This plant has a single flower stem and grows from 1.5-16 inches in wet locations at 4,000-11,000 feet.

California wildflower identification photograph. Sierra Gentian, Gentianopsis holopetala.

Sierra Gentian

A similar but higher-altitude species, 5,000-13,000 feet, the Sierra Gentian (Gentianopsis holopetala) also grows in wet meadows and blooms in late summer. The petal lobes of Sierra Gentian are broader and shorter than Hikers Fringed Gentian. Sierra Gentian’s sepals are green, often with a central maroon mid-rib. The lower portion of the corolla is white, with purple longitudinal lines extending from the purple petal lobes. A plant may have one to several stems.

Sierra Gentian, Gentianopsis holopetala, California wildflowers playing card.

Sierra Gentian playing card, Deck B

California wildflower identification photograph. Dwarf Gentian, Gentianella amarella.

Autumn Dwarf Gentian

Unlike the other small, late-blooming meadow gentians presented, Autumn Dwarf Gentian (Gentianella amarella) has multiple flowers per stalk. Its flowers are rose-violet, blue, or white, with a distinctive fringe on the inner petals. The plant is two-32 inches tall and grows at 5,000-11,500 feet. Autumn Dwarf Gentian’s distribution includes both North America and Eurasia. 


America’s Peonies

California wildflower identification photograph. Brown’s Peony, Paeonia brownii.

Brown's Peony is one of only two peonies native to the Americas

 

Only two peony species are native to the Americas, and both are found in California. California Peony (Paeonia californica) is restricted to the coastal mountains of southern California and Baja California, while Brown’s Peony (Paeonia brownii) has a broader distribution growing in eight western U.S. states. In California, Brown’s peony is found in northern mountain regions. You will likely only see these plants in the wild, as both species are challenging to cultivate.

Sierra Nevada wildflower, Brown’s Peony, Paeonia brownii.

Leaves of Brown's Peony

Brown’s Peony grows about one foot tall. Its leaves are thick and grayish blue-green, with multiple divisions. It is easiest to locate the flowers by first recognizing the plant. Flowers droop and may be hidden by leaves. When flowers are visible, the sepals frequently block other flower structures from an overhead view.

Sierra Nevada photography. Brown’s Peony, Paeonia brownii.

Flower of Brown's Peony

The one to two-inch flower of Brown’s Peony is distinctive and unforgettable. Both the sepals and petals are thick. A flower typically has five green or maroon sepals. The petals are maroon to red and yellow at the edges. Petals range in number from five to ten. Yellow stamens contrast nicely with the reddish petals. Stamens are variable in number and numerous; nearly 100 may be present. Two to five large carpels are yellow-green, turning to orange at maturity.

Brown’s Peony, Paeonia brownii, California wildflowers playing card.

Brown's Peony Playing Card, Deck A


SIERRA'S SHOOTING STARS

Henderson’s Shooting Star, Primula hendersonii, flowers shoot up as they mature.

shootinG star flowers change positions during development

Shooting Stars in the genus Primula (family Primulaceae) are among the most recognizable plants in the Sierra. Leaves grow at the base of the plants; leafless flower stalks rise above, producing dart-like flowers. Petals, fused at the base, are swept backwards with a wind-blown appearance, while the pistil and stamens protrude forward. Sepals are hidden under the reflexed petals. Young flowers face down.  After pollination, the flowers “shoot” upward with their stigmas pointing to the sky. 

Flower structure: Henderson’s Shooting Star, Primula hendersonii, labeled.

Shooting star flower morphology

Shooting stars have radial symmetry, a superior ovary, and four or five petals, stamens, and fused sepals. Petals may be white, pink, or magenta. The base of the petals is frequently yellow, with additional maroon and white markings. Stamen filaments may be free or fused to form a tube. The anthers’ connective tissue may be smooth or wrinkled and a single color or bicolored. The stigma may be bulbous or not. Noting these features is essential in identifying individual shooting star species.

California wildflower identification photograph. Padre’s Shooting Star, Primula clevelandii patula,

Padre’s Shooting Star

Eight species of shooting stars are native to California, with seven species found in the Sierra Nevada. Padre’s Shooting Star, Primula clevelandii patula, is a California endemic plant with a distribution that includes the western Sierra foothills, reaching elevations of up to 2,000 feet. Padre’s Shooting Star has spoon-shaped leaves, more than two times longer than wide. The plant is an early bloomer (March-May) that prefers seasonally wet locations in serpentine or alkaline soils.

Padre’s Shooting Star, Primula clevelandii patula, flower detail.

Padre’s Shooting Star's flower

Padre’s Shooting Star’s flower stalk may grow to 18 inches, producing one to six flowers; the petals may be white or pink. The filament tube is broad, maroon to almost black in color, often with white or yellow tips. Anthers are blue to purple and rounded or notched at the tip. The stigma is the same width as the style.

California wildflower identification photograph. Henderson’s Shooting Star, Primula hendersonii.

Henderson’s Shooting Star

Henderson’s Shooting Star, Primula hendersonii, resembles Padre’s Shooting Star; their distribution ranges overlap, and they may hybridize. Henderson’s Shooting Star grows in partial shade, in drier locations than Padre’s Shooting Star, and has a greater elevation range of up to 6,200 feet. Henderson’s Shooting Star has rounder leaves, with a length less than two times the width.

Henderson’s Shooting Star, Primula hendersonii, bulblets.

Henderson’s Shooting Star produces cloned colonies

Whereas Padre’s Shooting Star plants are often solitary, Henderson’s Shooting Stars are more likely to grow in crowded populations. This is because Henderson’s Shooting Star plants produce tiny bulblets (insert) at their stem/root interface. These bulblets allow the plants to reproduce nearby clones vegetatively.

Sierra Nevada photography: Henderson’s Shooting Star, Primula hendersonii.

Henderson's shooting star's flower

Henderson’s Shooting Star is a spring bloomer producing from three to 17 pink, magenta, lavender, or occasionally white-petaled flowers per stalk. The filament connectives are wrinkled and a solid dark magenta to black, with no white spots. Henderson’s Shooting Star has stigmas the same width as the style. The stamen/pistil region is narrow and resembles a mosquito proboscis, giving the plant its second common name, Mosquito Bill.

California wildflower identification photograph. Sierra Shooting Star, Primula jeffreyi.

Jeffrey’s/Sierra Shooting Star

Jeffrey’s or Sierra Shooting Star, Primula jeffreyi, is a late spring/summer bloomer found in meadows and on streambanks at elevations from 2,000 to 9,800 feet. It is a larger plant than the previous shooting stars, growing up to two feet tall, and has distinctly different leaves, which are larger, measuring up to 20 inches in length, and linear or elongated. 

Sierra Shooting Star, Primula jeffreyi, flowers.

Jeffrey’s/Sierra Shooting Star's flowers

Although the flower structure of Jeffrey’s Shooting Star appears similar to the species mentioned above, there are some distinct differences. Jeffrey’s Shooting Star has a larger number of flowers per flower stalk, up to 18. Stamen filaments are free or only partially fused. The pistil has a slightly enlarged stigma, and glandular hairs are present on the stem and flowers. A magnifying glass may be required to see the enlarged stigma and glandular hairs.

Sierra Shooting Star, Primula jeffreyi, California wildflowers playing cards.

Sierra shooting star Card, Deck B

California wildflower identification photograph. Alpine Shooting Star, Primula tetrandra.

Alpine Shooting Star

Alpine Shooting Star, Primula tetrandra, looks like a smaller version of Jeffrey’s Shooting Star. The stem grows to only 12 inches, and the leaves are eight inches or less in length. Alpine Shooting Star is a summer-blooming species found in moist subalpine and alpine locations at elevations up to 11,000 feet.  

Alpine Shooting Star, Primula tetrandra, flower detail.

Alpine Shooting Star has an enlarged stigma

Alpine Shooting Star has a stem lacking glandular hairs, stamen filaments are entirely free, and the stigma is notably broader than the style. Note the enlarged stigmas at the arrows in the photo.


Other Sierra Shooting Star species are less widely distributed. They include Scented Shooting Star (Primula fragrans), Beautiful Shooting Star (Primula pauciflora), and Subalpine Shooting Star (Primula subalpina). 

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